All this is the necessary consequence of the process of development.
For the earthy matter in the body issuing from it is converted into parts that are useful as weapons. That which flows upwards gives hardness or size to the beak; and, should any flow downwards, it either forms spurs upon the legs or gives size and strength to the claws upon the feet. But it does not at one and the same time produce both these results, one in the legs, the other in the claws; for such a dispersion of this residual matter would destroy all its efficiency. In other birds this earthy residue furnishes the legs with the material for their elongation; or sometimes, in place of this, fills up the interspaces between the toes. Thus it is simply a matter of necessity, that such birds as swim shall either be actually web-footed, or shall have a kind of broad blade-like margin running along the whole length of each distinct toe. The forms, then, of these feet are simply the necessary results of the causes that have been mentioned. Yet at the same time they are intended for the animal's advantage. For they are in harmony with the mode of life of these birds, who, living on the water, where their wings are useless, require that their feet shall be such as to serve in swimming. For these feet are so developed as to resemble the oars of a boat, or the fins of a fish; and the destruction of the foot-web has the same effect as the destruction of the fins; that is to say, it puts an end to all power of swimming.